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Welles Papers, Series 5: Field Notes, Box 10–11, University of California Museum of Paleontology Archives). Kerley introduced Welles to a Navajo man named Jesse Williams who had discovered the remains of a large dinosaur in 1940 near the now-famous dinosaur trackway site at Moenave, Arizona (Field Notes Summer of 1942 May 12–July 19 Arizona, The Samuel P. Camp returned to Berkeley, California and left Welles to meet with a trader named Richard Kerley at Tuba City, Arizona. Camp was primarily interested in the Permian rocks near Monument Valley, and Welles spent most of the summer in the Moenkopi Formation near Cameron, Arizona. The summer of 1942 began with Sam Welles assisting Charles Camp of the University of California Museum of Paleontology in his annual field season in the American Southwest. Many characters of the axial skeleton of Dilophosaurus wetherilli are derived compared to Late Triassic theropods and may be associated with macropredation and an increase in body size in Theropoda across the Triassic-Jurassic boundary. We did not recover a monophyletic ‘Dilophosauridae.’ Instead of being apomorphic for a small clade of early theropods, it is more likely that elaboration of the nasals and lacrimals of stem-averostrans is plesiomorphically present in early ceratosaurs and tetanurans that share those features. Dilophosaurus wetherilli is not recovered as a ceratosaur or coelophysoid, but is instead a non-averostran neotheropod in a grade with other stem-averostrans such as Cryolophosaurus ellioti and Zupaysaurus rougieri. When all five specimens are included in an analysis, they form a monophyletic clade that supports the monotypy of the genus.

Iterative specimen-based phylogenetic analyses result in each of the additional specimens recovered as the sister taxon to the holotype.

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The cervical vertebrae exhibit serial variation within the posterior centrodiapophyseal lamina, which bifurcates and reunites down the neck. The parasagittal nasolacrimal crests are uniquely constructed by a small ridge on the nasal process of the premaxilla, dorsoventrally expanded nasal, and tall lacrimal that includes a posterior process behind the eye. Redescription of the holotype, referred, and previously undescribed specimens of Dilophosaurus wetherilli supports the existence of a single species of crested, large-bodied theropod in the Kayenta Formation.

Skeletons of this species were collected from the middle and lower part of the Kayenta Formation in the Navajo Nation in northern Arizona. Despite its charismatic presence in pop culture and dinosaurian phylogenetic analyses, major aspects of the skeletal anatomy, taxonomy, ontogeny, and evolutionary relationships of this dinosaur remain unknown. Dilophosaurus wetherilli was the largest animal known to have lived on land in North America during the Early Jurassic.

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